• Aavik, T., Carmona, C.P., Träger, S., Kaldra, M., Reinula, I., Conti, E., Keller, B., Helm, A., Hiiesalu, I., Hool, K., Kaisel, M. et al. (2020). Landscape context and plant population size affect morph frequencies in heterostylous Primula veris‐results of a nationwide citizen‐science campaign. Journal of Ecology, 108(6): 2169-2183.
  • Carmona, C.P., Guerrero, I., Peco, B., Morales, M.B., Oñate, J.J., Pärt, T., Tscharntke, T., Liira, J., Aavik, T., Emmerson, M., Berendse, F., (2020). Agriculture intensification reduces plant taxonomic and functional diversity across European arable systems. Functional Ecology, 34(7): 1448-1460.
  • Hannula, SE, Träger, S (2020). Soil fungal guilds as important drivers of the plant richness-productivity relationship. New Phytologist, 226: 947-949. doi:10.1111/nph.16523.
  • Alahuhta, J., Antikainen, H., Hjort, J., Helm, A. & Heino, J., (2020) Current climate overrides historical effects on species richness and range size of freshwater plants in Europe and North America. Journal of Ecology.
  • Smith, A.L., Hodkinson, T.R., Villellas, J., Catford, J.A., Csergő, A.M., Blomberg, S.P., Crone, E.E., Ehrlén, J., Garcia, M.B., Laine, A.L., Roach, D.A., … Helm, A. et al. (2020) Global gene flow releases invasive plants from environmental constraints on genetic diversity. Proceedings of the National Academy of Sciences.
  • Kattge, J., Bönisch, G., Díaz, S., ... Helm, A., ... et al. (2020) TRY plant trait database – enhanced coverage and open access. Global Change Biology, 26: 119–188.
  • Blaus, A.; Reitalu, T.; Gerhold, P.; Hiiesalu, I.; Massante, J.C.; Veski, S. (2020). Modern pollen–plant diversity relationships inform palaeoecological reconstructions of functional and phylogenetic diversity in calcareous fens. Frontiers in Ecology and Evolution, 8, 207. DOI: 10.3389/fevo.2020.00207.


  • Massante, J.C., Götzenberger, L., Takkis, K., Hallikma, T., Kaasik, A., Laanisto, L., Hutchings, M.J., Gerhold, P. 2019. Publisher Correction: Contrasting latitudinal patters in phylogenetic diversity between woody and herbaceous communities. Scientific Reports 9: 7917.
  • Massante, J.C., Götzenberger, L., Takkis, K., Hallikma, T., Kaasik, A., Laanisto, L., Hutchings, M.J., Gerhold, P. 2019. Contrasting latitudinal patters in phylogenetic diversity between woody and herbaceous communities. Scientific Reports 9: 6443.
  • Carvalheiro, L.G., Biesmeijer, J.C., Franzén, M., Aguirre-Gutierrez, J., Garibaldi, L.A., Helm, A., Michez, D., Pöyry, J., Reemer, M., Schweiger, S., Berg, L. van den, WallisDeVries, M.F., Kunin., W.E., 2019. Nitrogen deposition shaped the composition of plant and pollinator assemblages during the past century. Ecography, 
  • Franzén, M., Dieker, P., Schrader, J., Helm, A. (2019). Rapid plant colonization of the forelands of a vanishing glacier is strongly associated with species traits. Arctic, Antarctic, and Alpine Research, 51, 366–378.
  • Noreika, N., Helm, A., Öpik, M., Jairus, T., Vasar, M., Reier, Ü., Kook, E.; Riibak, K., Kasari, L., Tullus, H.,Tullus, T., Lutter, R., Oja, E., Saag, A., Randlane, T., Pärtel, M. (2019). Forest biomass, soil and biodiversity relationships originate from biogeographic affinity and direct ecological effects. Oikos, 128, 1653-1665.
  • Aavik, T., Thetloff, M., Träger, S., Hernández-Agramonte, I. M., Reinula, I., & Pärtel, M. (2019). Delayed and immediate effects of habitat loss on the genetic diversity of the grassland plant Trifolium montanum. Biodiversity and Conservation, 1-21.
  • Reitalu, T., Bjune, A. E., Blaus, A., Giesecke, T., Helm, A., Matthias, I., Peglar, S. S., Salonen, S. J., Seppä, H., Väli, V., Birks, H. J. B. (2019). Patterns of modern pollen and plant richness across northern Europe. Journal of Ecology, 107, 1662-1677.
  • Plue, J., Aavik, T., Cousins, S. A. O. (2019). Grazing networks promote plant functional connectivity among isolated grassland communities. Diversity and Distributions, 25: 102-115.
  • Träger, S., Öpik, M., Vasar, M., Wilson, S. D. (2019). Belowground plant parts are crucial for comprehensively estimating total plant richness in herbaceous and woody habitats. Ecology, 100: e02575.


  • Takkis, K., Kull, T., Hallikma, T., Jaksi, P., Kaljund, K., Kauer, K., Kull, T., Kurina, O., Külvik, M., Lanno, K., Leht, M., Liira, J., Melts, I., Pehlak, H., Raet, J., Sammet, K., Sepp, K., Väli, Ü., Laanisto, L. 2018. Drivers of species richness and community integrity of small forest patches in an agricultural landscape. Journal of Vegetation Science 29(6): 978-988.   
  • Takkis, K., Tscheulin, T., Petanidou, T. 2018. Differential effects of climate warming in the nectar secretion of early- and late-flowering Mediterranean plants. Frontiers in Plant Science 8: 874.
  • Aavik, T. & Helm, A. (2018). Restoration of plant species and genetic diversity depends on landscape-scale dispersal. Restoration Ecology, 26:S92-S102.
  • Török, P., Helm, A., Kiehl, K., Buisson, E., Valkó, O. (2018). Beyond the species pool: modification of species dispersal, establishment, and assembly by habitat restoration. Restoration Ecology, 26:S65-S72.
  • Van Geel, M., Ceulemans, T., Plue, J., Takkis, K., Saar, L., Kasari, L., Peeters, G., van Acker, K., Jacquemyn, H. & Honnay, O. 2018. Environmental filtering rather than host plant specificity shapes the arbuscular mycorrhizal communities of European semi-natural grasslands. New Phytologist 220: 1262–1272.
  • Elbakidze, M., Hahn, T., Dawson, L., Zimmermann, N.E., Cudlín, P., Friberg, N., Genovesi, P., Guarino, R., Helm, A., Jonsson, B. and Lengyel, S., Leroy, B., Luzzati, T., Milbau, A., Pérez-Ruzafa, A., Roche, P., Roy, H., Sabyrbekov, R., Vanbergen, A., Vandvik, V. (2018). Direct and indirect drivers of change in biodiversity and nature’s contributions to people. In: M. Rounsevell, M. Fischer, A. Torre-Marin Rando, A. Mader (Ed.). The IPBES regional assessment report on biodiversity and ecosystem services for Europe and Central Asia (385−568). Bonn, Germany: Secretariat of the Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services.
  • Dengler, J., Wagner, V., Dembicz, I., Garcia-Mijangos, I., Naqinezhad, A., Boch, S., Chiarucci, A., Conradi, T., Filibeck, G., Guarino, R., Janisova, M., Steinbauer, M. J., Acic, S., Acosta, A. T. R., ... Helm, A., ... Biurrun, I. (2018). GrassPlot - a database of multi-scale plant diversity in Palaearctic grasslands. Phytocoenologia, 48 (3), 331−347.10.1127/phyto/2018/0267.
  • Tyler, T.; Bengtsson, F.; Dahlberg, C. J.; Lönnell, N.; Hallingbäck, T.; Reitalu, T. (2018). Determinants of bryophyte species composition and diversity on the Great Alvar of Öland, Sweden. Journal of Bryology, 40 (1), 12−30. DOI: 10.1080/03736687.2017.1412387.


  • Aavik, T., Talve, T., Thetloff, M., Uuemaa, F., Oja, T. (2017). Genetic consequences of landscape change for rare endemic plants - a case study of Rhinanthus osiliensis. Biological Conservation, 210: 125-135.
  • Dengler, J., Wagner, V., Dembicz, I., García-Mijangos, I., Naqinezhad, A., Boch, S., ... Helm, A., ... & Janisova, M. (2018). GrassPlot-a database of multi-scale plant diversity in Palaearctic grasslands. Phytocoenologia, 48(3), 331-347.
  • Saar, L., de Bello, F., Pärtel, M., Helm, A. (2017). Trait assembly in grasslands depends on habitat history and spatial scale. Oecologia, 184, 1−12.
  • Bueno, C.G., Moora, M., Gerz, M., Davison, J., Öpik, M., Pärtel, M., Helm, A., Ronk, A., Kühn, I., Zobel, Z. (2017). Plant mycorrhizal status, but not type, shifts with latitude and elevation in Europe. Global Ecology and Biogeography, 26 (6), 690−699.
  • Randlane, T., Tullus, T., Saag, A., Lutter, R., Tullus, A., Helm, A., Tullus, H., Pärtel, M. (2017). Diversity of lichens and bryophytes in hybrid aspen plantations in Estonia depends on landscape structure. Canadian Journal of Forest Research , 47, 1202−1214.
  • Török, P., Helm, A. (2017). Ecological theory provides strong support for habitat restoration. Biological Conservation, 206, 85−91.
  • Träger, S., Milbau, A., Wilson, S. D. (2017). Potential contributions of root decomposition to the nitrogen cycle in arctic forest and tundra. Ecology and Evolution, 7: 11021-11032.
  • Träger, S., Wilson, S. D. (2017). Root heterogeneity along an arctic elevational gradient: the importance of resolution. Functional Ecology, 31: 480-487.
  • Price, J., Tamme, R., Gazol, A., de Bello, F., Takkis, K., Uria Diez, J., Kasari, L. & Pärtel, M. 2017. Within-community environmental variability drives trait variability in species-rich grasslands. Journal of Vegetation Science 28: 303–312.
  • Lewis, R.J., de Bello, F., Bennett, J.A., Fibich, P., Finerty, G.E., Götzenberger, L., Hiiesalu, I., Kasari, L., Lepš, J., Májeková, M., Mudrák, O., Riibak, K., Ronk, A., Rychtecká, T., Vitová, A. & Pärtel, M. 2017. Applying the dark diversity concept to nature conservation. Conservation Biology 31: 40–47.

2016 and earlier

  • Kasari, L., Saar, L., de Bello, F., Takkis, K., Helm, A. (2016). Hybrid ecosystems can contribute to local biodiversity conservation. Biodiversity and Conservation, 25 (14), 3023-3041.
  • Kook, E., Pihu, S., Reier, Ü., Thetloff, M., Aavik, T., Helm, A. (2016). Do landscape dissimilarity and environmental factors affect genetic and phenotypic variability in Myosotis laxa s. lato (Boraginaceae)? Annales Botanici Fennici, 53:56−66.
  • Meriste, M., Helm, A., Ivask, M. (2016). Ground-dwelling spider fauna of flooded meadows in Matsalu, Estonia. Wetlands, 36, 525−537.
  • Emmerson, M., Morales, M. B., Oñate, J. J., Batáry, P., Berendse, F., Liira, J., Aavik, T., Guerrero, I., Bommarco, R., Eggers, S., Pärt, T., Tscharntke, T., Weisser, W., Clement, L. & Bengtsson, J. (2016). How agricultural intensification affects biodiversity and ecosystem services. Advances in Ecological Research, 55: 43−97.
  • Májeková, M., Paal, T., Plowman, N.S., Bryndová, M., Kasari, L., Norberg, A., Weiss, M., Bishop, T.R., Luke, S.H., Sam, K., Le Bagousse-Pinguet, Y., Lepš, J., Götzenberger, L. & de Bello, F. 2016. Evaluating functional diversity: Missing trait data and the importance of species abundance structure and data transformation. PLoSOne 11: e0152532.
  • Takkis, K., Tscheulin, T., Tsalkatis, P., Petanidou, T. 2015. Climate change reduces nectar secretion in two common Mediterranean plants. Annals of Botany PLANTS 7: plv111.
  • Dallimer, M., Jacobsen, J.B., Lundhede, T.H., Takkis, K., Giergiczny, M., Thorsen, B.J. 2015. Patriotic values for public goods: transnational trade-offs for biodiversity and ecosystem services. BioScience 65(1): 33-42. 
  • Helm, A. (2015). Habitat restoration requires landscape-scale planning. Applied Vegetation Science, 18 (2), 177−178.
  • Bagaria, G., Helm, A., Rodà, F., Pino, J. (2015). Assessing coexisting plant extinction debt and colonization credit in a grassland–forest change gradient. Oecologia, 179 (3), 823−834.
  • Helm, A., Zobel, M., Moles, A.T., Szava-Kovats, R., Pärtel, M. (2015). Characteristic and derived diversity: implementing the species pool concept to quantify conservation condition of habitats. Diversity and Distributions, 21 (6), 711−721.
  • Aavik, T., Holderegger, R., Bolliger, J. (2014). The structural and functional connectivity of the grassland plant Lychnis flos-cuculi. Heredity, 112: 471 - 478.
  • Aavik, T., Bosshard, D., Edwards, P.J., Holderegger, R., Billeter, R. (2014). Fitness in naturally occurring and restored populations of a grassland plant Lychnis flos-cuculi in a Swiss agricultural landscape. Restoration Ecology, 22: 98 - 10.
  • Helm, A., Kalamees, R., Zobel, M. (2014). Vegetation patterns and their underlying processes: Where are we now? Journal of Vegetation Science, 25(5), 1113−1116.
  • Sutcliffe, L.M.E.,Batary, P., Kormann, U., Baldi, A., Dicks, L.V., Herzon, I., Kleijn, D., Tryjanowski, P., Apostolova, I., Arlettaz, R., Aunins A., Aviron, S., Baležentiene, L., Fischer, C., Halada, L., Hartel, T., Helm, A., Hristov, I., Jelaska, S.D., Kaligaric, M., Kamp, J., Klimek, S., Koorberg, P., Kostiukova, J., Kovacs-Hostyanszki, A., Kuemmerle, T., Leuschner, C., Lindborg, R., Loos, J., Maccherini, S., Marja, R., Mathe, O., Paulini, I., Proenca, V., Rey-Benayas, J., Sans, F.X., Seifert, C., Stalenga, J., Timaeus, J., Torok, P., van Swaay, C., Viik, E., Tscharntke, T. (2014). Harnessing the biodiversity value of Central and Eastern European farmland. Diversity and Distributions, 21, 722–730.
  • Riibak, K., Reitalu, T., Tamme, R., Helm, A., Gerhold, P., Znamenskiy, S., Bengtsson, K., Rosén, E., Prentice, H., Pärtel, M. (2015). Dark diversity in dry calcareous grasslands is determined by dispersal ability and stress-tolerance. Ecogaphy, 38, 713−721.
  • Aavik, T., Bosshard, D., Edwards, P., Holderegger, R., Billeter, R. (2014). Effets de mélanges de semences sur la diversité génétique et la performance. Recherche Agronomique Suisse, 5: 20 - 27.
  • Aavik, T., Bosshard, D., Edwards, P., Holderegger, R. & Billeter, R. 2014. Genetische Vielfalt in Wildpflanzen-Samenmischungen. Agrarforschung Schweiz, 5: 20 - 27.
  • Aavik, T., Holderegger, R., Edwards, P.J. & Billeter, R. (2013). Patterns of contemporary gene flow suggest low functional connectivity of grasslands in a fragmented agricultural landscape. Journal of Applied Ecology, 50: 395 - 403.
  • Seabloom, E. W., E. T. Borer ,... Helm, A., ... Yang, L. (2013). Predicting invasion in grassland ecosystems: is exotic dominance the real embarrassment of richness? Global Change Biology 19: 3677−3687.
  • Kasari, L., Gazol, A., Kalwij, J.M., Helm, A. (2013). Low shrub cover in alvar grasslands increases small-scale diversity by promoting the occurrence of generalist species. Tuexenia, 33, 293−308.
  • Takkis, K., Pärtel, M., Saar, L. & Helm, A. (2013). Extinction debt in a common grassland species: immediate and delayed responses of plant and population fitness. Plant Ecology, 214, 953−963.
  • Reitalu, T., Helm, A., Pärtel, M., Bengtsson, K., Gerhold, P., Rosén, E., Takkis, K., Znamenskiy, S., Prentice, H. C. (2014). Determinants of fine-scale plant diversity in dry calcareous grasslands within the Baltic Sea region. Agriculture, Ecosystems & Environment, 182, 59−68.
  • Gazol, A., Tamme, R., Takkis, K., Kasari, L., Saar, L., Helm, A., Pärtel, M. (2012). Landscape-and small-scale determinants of grassland species diversity: direct and indirect influences. Ecography, 35, 944−951.
  • Guerrero, I., Morales, M.L.B., Oñate, J.J., Geiger, F., Berendse, F., Snoo, G. de, Eggers, S., Pärt, T., Bengtsson, J., Clement, L.W., Weisser, W.W., Olszewski, A., Ceryngier, P., Hawro, V., Liira, J., Aavik, T., Fischer, C., Thies, C., Tscharntke, T. (2012). Response of ground-nesting farmland birds to agricultural intensification across Europe: landscape and field level management factors. Biological Conservation, 152: 74 - 80.
  • Lindborg, R., Helm, A., Bommarco, R., Heikkinen, R. K., Kühn, I., Pykälä, J., Pärtel, M. (2012). Effect of habitat area and isolation on plant trait distribution in European forests and grasslands. Ecography, 35, 356−363.
  • Aavik, T., Edwards, J.P., Holderegger, R., Graf, R., Billeter, R. (2012). Genetic consequences of using seed mixtures in restoration: a case study of a wetland plant Lychnis flos-cuculi. Biological Conservation, 145: 195 - 204.
  • Marini, L., Bruun, H. H., Heikkinen, R. K., Helm, A., Honnay, O., Krauss, J., Kühn, I., Lindborg, R., Pärtel, M., Bommarco, R. (2012). Traits related to species persistence and dispersal explain changes in plant communities subjected to habitat loss. Diversity and Distributions, 18, 898−908.
  • Saar, L., Takkis, K., Pärtel, M., Helm, A. (2012). Which plant traits predict species loss in calcareous grasslands with extinction debt? Diversity and Distributions, 18, 808−817.
  • Moles, A.T., Flores-Moreno, H., Bonser, S.P., Warton, D.I., Helm, A., Warman, L., Eldridge, D.J., Jurado, E., Hemmings, F.A., Reich, P.B., Cavender-Bares, J., Seabloom, E.W., Mayfield, M.M., Sheil, D., Djietror, J.C., Peri, P.L., Enrico, L., Cabido, M.R., Setterfield, S.A., Lehmann, C.E.R., Thomson, F.J. (2012). Invasions: the trail behind, the path ahead, and a test of a disturbing idea. Journal of Ecology, 100, 116−127.
  • Guerrero, I., Morales, M., Onante, J.J., Aavik, T., Bengtsson, J., Berendse, F., Clement, L., Dennis, C., Eggers, S., Emmerson, M., Fischer, C., Flohre, A., Geiger, F., Hawro, V., Inchausti, P., Kalamees, A., Kinks, R., Liira, J., Melendez, L., Pärt, T., Thies, C., Tscharntke, T., Olszewski, A., Weisser, W. (2011). Taxonomic and functional diversity of farmland bird communities across Europe: effects of biogeography and agricultural intensification. Biodiversity and Conservation, 20: 3663 - 3681.
  • Flohre, A., Fischer, C., Aavik, T., Bengtsson, J., Berendse, F., Bommarco, R., Ceryngier, P., Clement, L.W., Dennis, C., Eggers, S., Emmerson, M., Geiger, F., Guerrero, I., Hawro, V., Inchausti, P., Liira, J., Morales, M.B., Oñate, J.J., Pärt, T., Weisser, W.W., Winqvist, C., Thies, C., Tscharntke, T. (2011). Agricultural intensification and biodiversity partitioning in European landscapes comparing plants, carabids, and birds. Ecological Applications, 21: 1772 - 1781.
  • Winqvist, C., Bengtsson, J., Aavik, T., Berendse, F., Clement, L. W., Eggers, S., Fischer, C., Flohre, A., Geiger, F., Liira, J., Pärt, T., Thies, C., Tscharntke, T., Weisser, W. W., Bommarco, R. (2011). Mixed effects of organic farming and landscape complexity on farmland biodiversity and biological control potential across Europe. Journal of Applied Ecology, 48: 570 - 579.
  • Krauss, J., Bommarco, R., Guardiola, M., Heikkinen, R., Helm, A., Kuussaari, M., Lindborg, R., Öckinger, E., Pärtel, M., Pino, J., Pöyry, J., Raatikainen, K., Sang, A., Stefanescu, C. Teder, T., Zobel, M., Steffan-Dewenter, I. (2010). Habitat fragmentation causes immediate and time-delayed biodiversity loss at different trophic levels. Ecology Letters, 13, 597–605.
  • Sang, A., Teder, T., Helm, A., Pärtel, M. (2010). Indirect evidence for an extinction debt of grassland butterflies half century after habitat loss. Biological Conservation, 143, 1405–1413.
  • Geiger, F., Bengtsson, J., Berendse, F., Weisser, Wolfgang W., Emmerson, M., Morales, M.B., Ceryngier, P., Liira, J., Tscharntke, T. Winqvist, C., Eggers, S., Bommarco, R., Pärt, T., Bretagnolle, V., Plantegenest, M., Clement, L.W., Dennis, C., Palmer, C., Onate, J.J., Guerrero, I., Hawro, V., Aavik, T., Thies, C., Flohre, A., Hänke, S., Fischer, C., Goedhart, P.W., Inchausti, P. (2010). Persistent negative effects of pesticides on biodiversity and biological control potential on European farmland. Basic and Applied Ecology, 11: 97 - 105.
  • Aavik, T., Liira, J. (2010). Quantifying the effect of organic farming, field boundary type and landscape structure on the vegetation of field boundaries. Agriculture, Ecosystems and Environment, 135: 178 - 186.
  • Helm, A., Oja, T., Saar, L., Takkis, K., Talve, T., Pärtel, M. (2009). Human influence lowers plant genetic diversity in communities with extinction debt. Journal of Ecology, 97, 1329−1336.
  • Kuussaari, M., Bommarco, R., Heikkinen, R.K., Helm, A., Krauss, J., Lindborg, R., Öckinger, E., Pärtel, M., Pino, J., Roda, F., Stefanescu, C., Teder, T., Zobel, M.,  Steffan-Dewenter, I. (2009). Extinction debt: a challenge for biodiversity conservation. Trends in Ecology & Evolution, 24, 564−571.
  • Aavik, T., Püssa, K., Roosaluste, E., Moora, M. (2009). Vegetation change in boreonemoral forest during succession – trends in species composition, richness and differentiation diversity. Annales Botanici Fennici, 46: 326-335.
  • Aavik, T., Liira, J. (2009). Agrotolerant and high nature-value species - plant biodiversity indicator groups in agroecosystems. Ecological Indicators, 9: 892-901.
  • Liira, J., Schmidt, T., Aavik, T., Arens, P., Augenstein, I., Bailey, D., Billeter, R., Bukacek, R., Burel, F., De Blust, G., De Cock, R., Dirksen, J., Edwards, P. J., Hamersky, R., Herzog, F., Klotz, S., Kühn, I., Le Coeur, D., Miklova, P., Roubalova, M., Schweiger, O., Smulders, M. J. M., van Wingerden,W. K. R. E., Bugter, R., Zobel, M. (2008). Plant functional group composition and large-scale species richness in the agricultural landscapes of Europe. Journal of Vegetation Science, 19: 3-14.
  • Aavik, T., Augenstein, I., Bailey, D., Herzog, F., Zobel, M., Liira, J. (2008). What is the role of local landscape structure in the vegetation composition of field boundaries? Applied Vegetation Science, 11: 375-386.
  • Aavik, T., Jõgar, Ü., Liira, J., Tulva, I., Zobel, M. (2008). Plant diversity in a calcareous wooded meadow - The significance of management continuity. Journal of Vegetation Science, 19: 475-484.
  • Liira, J., Aavik, T., Parrest, O., Zobel, M. (2008). Agricultural sector, rural environment and biodiversity in the central and eastern European EU member states. Acta Geographica Debrecina Landscape and Environment Series, 2: 46-64.
  • Pärtel, M. & Helm, A. (2007). Invasion of woody species into temperate grasslands: Relationship with abiotic and biotic soil resource heterogeneity. Journal of Vegetation Science 18.1, 63−70
  • Pärtel, M., Helm, A., Reitalu, T., Liira, J., Zobel, M. (2007). Grassland diversity related to the Late Iron Age human population density. Journal of Ecology, 95, 574−582.
  • Znamenskiy, S., Helm, A. & Pärtel, M. (2006) Threatened alvar grasslands in NW Russia and their relationship to alvars in Estonia. Biodiversity and Conservation 15, 1797−1809.
  • Helm, A., Hanski, I., & Pärtel, M. (2006). Slow response of plant species richness to habitat loss and fragmentation. Ecology Letters, 9, 72−77.
  • Pärtel, M., Helm, A., Ingerpuu, N., Reier, Ü., Tuvi, E.-L. (2004). Conservation of Northern European plant diversity: the correspondence with soil pH. Biological Conservation 120, 525−531.
  • Dengler, J., Rusina, S., Boch, S., Bruun, H.H., Diekmann, M., Dierßen, K., Dolnik, C., Dupré, C., Golub, V.B., Grytnes, J.-A., Helm, A., Ingerpuu, N., Löbel, S., Pärtel, M., Rašomavicius, V., Tyler, G., Znamenskiy, S.R., Zobel, M. (2006) Working group on dry grasslands in the Nordic and Baltic region - Outline of the project and first results for the class Festuco-Brometea. Annali di Botanica Nuova Serie, 6: 128, Rome.
  • Helm, A., Urbas, P., Pärtel, M. (2007) Plant diversity and species characteristics of alvar grasslands in Estonia and Sweden. van der Maarel, E. (Ed.). Acta Phytogeographica Suecica, 88, 3342. Opulus Press.

Other scientific publications

  • Aavik, T., Luhamaa, H. & Kukk, T. (2009). Grasslands of Läänemaa County, Estonia. Veen, P., Jefferson, R., de Smidt, J., van der Straaten, J. (Ed-s.). In: Grasslands in Europe of high nature value. Zeist, the Netherlands: KNNV Publishing.
  • Pärtel, M., Helm, A., Roosaluste, E., Zobel, M. (2007) Bioloogiline mitmekesisus Eesti pool-looduslikes ökosüsteemides. Punning, J.M. (Ed.). Keskkonnauuringute nüüdisprobleeme (223 - 302). Tallinn: Tallinna Ülikooli Ökoloogia Instituut.
  • Bommarco, R., Öckinger, E., Helm, A. (2009) Life History Traits in Insects and Habitat Fragmentation. Atlas of Biodiversity Risk. Settele, J., Penev, L., Georgiev, T., Grabaum, R., Grobelnik, V., Hammen, V., Klotz, S., & Kühn, I. (Ed.) Pensoft Publishers, Sofia-Moscow. ISBN 978-954-642-446-4 (print) and ISBN 978-954-642-447-1.
  • Rosche, C, Schrieber, K, Hirsch, H, Blachnik, T, Träger, S, Seidler, G, Hensen, I (2014). Sexuelle Reproduktion und klonales Wachstum in kleinen Population von Antennaria dioica (L.) GAERTNER. Hercynia N.F., 47:59-86.